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  1. From Fish to Terrestrial Vertebrates

  One of the most significant evolutionary events that occurred on Earth was the transition of water-dwelling fish to terrestrial tetrapods (four-limbed organisms with backbones). Fish probably originated in the oceans, and our first records of them are in marine rocks. However, by the Devonian Period (408 million to 362 million years ago), they had radiated into almost all available aquatic habitats, including freshwater settings. One of the groups whose fossils are especially common in rocks deposited in fresh water is the lobe-finned fish.

  The freshwater Devonian lobe-finned fish rhipidistian crossopterygian is of particular interest to biologists studying tetrapod evolution. These fish lived in river channels and lakes on large deltas. The delta rocks in which these fossils are found are commonly red due to oxidized iron minerals, indicating that the deltas formed in a climate that had alternate wet and dry periods. If there were periods of drought, any adaptations allowing the fish to survive the dry conditions would have been advantageous. In these rhipidistians，several such adaptations existed. It is known that they had lungs as well as gills for breathing. Cross sections cut through some of the fossils reveal that the mud filling the interior of the carcass differed in consistency and texture depending on its location inside the fish. These differences suggest a sadlike cavity below the front end of the gut that can only be interpreted as a lung. Gills were undoubtedly the main source of oxygen for these fish, but the lungs served as an auxiliary breathing device for gulping air when the water became oxygen depleted, such as during extended periods of drought. So, these fish had already evolved one of the prime requisites for living on land: the ability to use air as a source of oxygen.

  A second adaptation of these fish was in the structure of the lobe fins. The fins were thick, fleshy, and quite sturdy, with a median axis of bone down the center. They could have been used as feeble locomotor devices on land, perhaps good enough to allow a fish to flop its way from one pool of water that was almost dry to an adjacent pond that had enough water and oxygen for survival. These fins eventually changed into short, stubby legs. The bones of the fins of a Devonian rhipidistian exactly match in number and position the limb bones of the earliest known tetrapods, the amphibians. It should be emphasized that the evolution of lungs and limbs was in no sense an anticipation of future life on land. These adaptations developed because they helped fish to survive in their existing aquatic environment.

  What ecological pressures might have caused fishes to gradually abandon their watery habitat and become increasingly land-dwelling creatures? Changes in climate during the Devonian may have had something to do with this if freshwater areas became progressively more restricted. Another impetus may have been new sources of food. The edges of ponds and streams surely had scattered dead fish and other water-dwelling creatures. In addition, plants had emerged into terrestrial habitats in areas near streams and ponds, and crabs and other arthropods were also members of this earliest terrestrial community. Thus, by the Devonian the land habitat marginal to freshwater was probably a rich source of protein that could be exploited by an animal that could easily climb out of water. Evidence from teeth suggests that these earliest tetrapods did not utilize land plants

  as food; they were presumably carnivorous and had not developed the ability to feed on plants.

  How did the first tetrapods make the transition to a terrestrial habitat? Like early land plants such as rhyniophytes, they made only a partial transition; they were still quite tied to water. However, many problems that faced early land plants were not applicable to the first tetrapods The ancestors of these animals already had a circulation system, and they were mobile, so that they could move to water to drink. Furthermore, they already had lungs, which rhipidistians presumably used for auxiliary breathing. The principal changes for the earliest tetrapods were in the skeletal system—changes in the bones of the fins, the vertebral column, pelvic girdle, and pectoral girdle.

  1. Paragraph 1 supports which of the following statements about fish evolution?

  A. Lobe-finned fish were among the earliest types of fish to appear.

  B. Fish began living in freshwater habitats only after originating elsewhere.

  C. Lobe-finned fish radiated into almost all available aquatic habitats.

  D. During the Devonian, lobe-finned fish were more common in marine than in freshwater habitats.

  2. According to paragraph 2, what do the minerals in the delta rocks containing rhipidistian crossopterygian fossils reveal?

  A. These deltas formed in dry periods but gradually became wetter.

  B.These deltas contain different types of iron minerals than do the surrounding areas.

  C.Most rhipidistian crossopterygian fish died when the climate became dry.

  D.Rhipidistian crossopterygian fish lived in areas that experienced alternate dry and wet periods.

  3. The word "advantageous" in the passage is closest in meaning to

  A. beneficial

  B. necessary

  C. remarkable

  D. common

  4. In paragraph 2, why does the author include the information that mud inside rhipidistian crossopterygian fossils differed in consistency and texture depending on where the mud was located?

  A. To provide evidence that rhipidistian crossopterygian lived in river channels and lakes on large deltas.

  B. To identify an effect of the oxidation of iron minerals on the evolution of rhipidistian crossopterygian.

  C. To help explain why scientists have concluded that rhipidistian crossopterygian probably had lungs.

  D. To explain why scientists decided to cut cross sections through some fossils of rhipidistian crossopterygian.

  5. Which of the sentences below best expresses the essential information in the highlighted sentence in the passage? Incorrect choices change the meaning in important ways or leave out

  essential information.

  A. Because the lungs of these fish were able to provide only a small amount of oxygen, these fish obtained most of their oxygen through their gills during periods of drought.

  B. During periods of extended drought, these fish used their lungs to increase their intake of oxygen beyond the levels absorbed by the gills in normal times.

  C. Although these fish primarily used their gills to obtain oxygen, they used their lungs to obtain oxygen from the air when there was not enough in the water.

  D. During periods of extended drought, the gills became an auxiliary breathing device and the lungs became the main source of oxygen for these fish.

  6. Which of the sentences below best expresses the essential information in the highlighted sentence in the passage? Incorrect choices change the meaning in important ways or leave out essential information.

  A. Because the lungs of these fish were able to provide only a small amount of oxygen, these fish obtained most of their oxygen through their gills during periods of drought.

  B. During periods of extended drought, these fish used their lungs to increase their intake of oxygen beyond the levels absorbed by the gills in normal times.

  C. Although these fish primarily used their gills to obtain oxygen, they used their lungs to obtain oxygen from the air when there was not enough in the water.

  D. During periods of extended drought, the gills became an auxiliary breathing device and the lungs became the main source of oxygen for these fish.

  7. The word "adjacent" in the passage is closest in meaning to:

  A. nearby

  B. available

  C. temporary

  D. fresh

  8. The word “progressively” in the passage is closest in meaning to:

  A. increasingly

  B. noticeably

  C. occasionally

  D. rapidly

  9. In paragraph 4, why does the author point out that crabs and other arthropods were already living on land when the ancestors of the first tetrapods began living there?

  A. To account for the presence of dead fish along the edges of ponds and streams during the Devonian.

  B. To support the claim that climate change caused freshwater habitats to become more restricted during the Devonian.

  C. To identify a consequence of the emergence of plants into terrestrial habitats near ponds and streams.

  D. To identify a possible reason for why certain fish gradually became terrestrial organisms.

  10. According to paragraph 4，teeth of the earliest tetrapods suggest that these tetrapods

  A. competed with other animals for protein

  B. were probably carnivores

  C. could easily climb out of water

  D. were able to eat plants

  11. According to paragraph 5, which of the following was true of the first tetrapods?

  A. They became dependent for food on organisms already living on land.

  B. They needed to develop new mechanisms for obtaining nutrients.

  C. They continued to live in close association with aquatic environments.

  D. They were evolutionarily far removed from their rhipidistian ancestors.

  12. According to paragraph 5, what was the main way that the earliest tetrapods differed from their immediate fish ancestors?

  A. The tetrapods had a different skeletal structure.

  B. The tetrapods had more sources of food available

  C. The tetrapods had a circulation system.

  D. The tetrapods could move to new pools of water.

  13. Look at the four squares [■] that indicate where the following sentence could be added to the passage.

  These would have been deposited by the receding waters of droughts, during which many aquatic animals must have died.

  Where would the sentence best fit? Click on a square [■] to add the sentence to the passage

  14. Directions: An introductory sentence for a brief summary of the passage is provided below. Complete the summary by selecting the THREE answer choices that express the most important ideas in the passage. Some sentences do not belong in the summary because they express ideas that are not presented in the passage or are minor ideas in the passage. This question is worth 2 points.

  Drag your answer choices to the spaces where they belong. To remove an answer choice, click on it.

  To review the passage, click VIEW TEXT

  Freshwater lobe-finned fish may be the direct ancestors of terrestrial tetrapods.

  A. Rhipidistian crossopterygian had features such as primitive lungs and thick fins that could have helped it survive dry periods.

  B. During the Devonian, the number of bones increased in the fins of rtiipidistians, improving such animals’ ability to swim and move over land

  C. Shortly after the earliest tetrapods developed lungs, plants and other animals began to flourish on land.

  D. By the Devonian period, lobe-finned fish preferred freshwater habitats to life in the ocean.

  E. A drier climate and new sources of food on land may have encouraged the lobe-finned fish’s

  move to a terrestrial existence.

  F. Early tetrapods remained closely connected to water, but several of their body structures were adapted for life on land.

  2. The Use of the Camera Obscura

  The precursor of the modern camera, the camera obscura is a darkened enclosure into which light is admitted through a lens in a small hole. The image of the illuminated area outside the enclosure is thrown upside down as if by magic onto a surface in the darkened enclosure. This technique was known as long ago as the fifth century B.C. in China. Aristotle also experimented with it in the fourth century B.C., and Leonardo da Vinci described it in his notebooks in 1490. In 1558 Giovanni Battista Della Porta wrote in his twenty-volume work Magia naturalis (meaning "natural magic") instructions for adding a convex lens to improve the quality of the image thrown against a canvas or panel in the darkened area where its outlines could be traced. Later, portable camera obscuras were developed, with interior mirrors and drawing tables on which the artist could trace the image. For the artist, this technique allows forms and linear perspective to be drawn precisely as they would be seen from a single viewpoint. Mirrors were also used to reverse the projected images to their original positions.

  Did some of the great masters of painting, then, trace their images using a camera obscura? Some ait historians are now looking for clues of artists' use of such devices. One of the artists whose paintings are being analyzed from this point of view is the great Dutch master, Jan Vermeer, who lived from 1632 to 1675 during the flowering of art and science in the Netherlands, including the science of optics. Vermeer produced only about 30 known paintings, including his famous The Art of Painting. The room shown in it closely resembles the room in other Vermeer paintings, with lighting coming from a window on the left, the same roof beams, and similar floor tiles, suggesting that the room was fitted with a camera obscura on the side in the foreground. The map hung on the opposite wall was a real map in Vermeers possession, reproduced in such faithful detail that some kind of tracery is suspected. When one of Vermeer’s paintings was X-rayed, it did not have any preliminary sketches on the canvas beneath the paint, but rather the complete image drawn in black and white without any trial sketches. Vermeer did not have any students, did not keep any records, and did not encourage anyone to visit his studio, facts that can be interpreted as protecting his secret use of a camera obscura.

  In recent times the British artist David Hockney has published his investigations into the secret use of the camera obscura, claiming that for up to 400 years, many of Western art’s great masters probably used the device to produce almost photographically realistic details in their paintings. He includes in this group Caravaggio, Hans Holbein, Leonardo da Vinci, Diego Velazquez,

  Jean-Auguste-Dominique Ingres, Agnolo Bronzino, and Jan van Eyck. From an artist’s point of view, Hockney observed that a camera obscura compresses the complicated forms of a three-dimensional scene into two-dimensional shapes that can easily be traced and also increases the contrast between light and dark, leading to the chiaroscuro effect seen in many of these paintings. In Jan van Eyck’s The Marriage of Giovanni Arnolfini and Giovanna Cenami, the complicated foreshortening in the chandelier and the intricate detail in the bride’s garments are among the clues that Hockney thinks point to the use of the camera obscura.

  So what are we to conclude? If these artists did use a camera obscura, does that diminish their stature? Hockney argues that the camera obscura does not replace artistic skill in drawing and painting. In experimenting with it, he found that it is actually quite difficult to use for drawing, and he speculates that the artists probably combined their observations from life with tracing of shapes.

  1. What can be inferred from paragraph 1 about Giovanni Battista Della Porta's contribution to the camera obscura?

  A. He translated a Chinese description of the use of the camera obscura and made the technique available to artists.

  B. His convex lens made the projected image easier to trace.

  C. His version of the camera obscura allowed for the later addition of a mirror.

  D. His improvements relied heavily on design changes proposed earlier by Leonardo da Vinci.

  2. The word “portable” in the passage is closest in meaning to

  A. valuable

  B. practical

  C. moveable

  D. popular

  3. The word "projected" in the passage is closest in meaning to

  A. whole

  B. corrected

  C. enlarged

  D. shown

  4. Paragraph 2 answers which of the following questions about paintings by Vermeer?

  A. What characteristics of Vermeer’s paintings suggest that he may have used a camera obscura?

  B. Why did Vermeer produce only about 30 paintings?

  C. Do Vermeer's paintings in general suggest that he was unable to paint accurately without using a camera obscura?

  D. Why did Vermeer need to draw an image on the canvas of the painting that was X-rayed if he was using a camera obscura?

  5. Which of the sentences below best expresses the essential information in the highlighted sentence in the passage? Incorrect choices change the meaning in important ways or leave out

  essential information.

  A. One artist with a particularly interesting point of view about the use of the camera obscura in painting was Jan Vermeer, who lived in the Netherlands from 1632 to 1675.

  B. Historical analyses suggest that Dutch masters were interested in the science of optics, so they may have used the camera obscura to trace images.

  C. The use of the camera obscura is being analyzed in the paintings of Jan Vermeer, who lived in the Netherlands when art and science were flourishing there.

  D. One view held by historians is that most Dutch masters were as interested in art as they were in science, and that provides clues about the techniques used in their paintings.

  6. What does paragraph 2 indicate about Vermeer’s The Art of Painting ?

  A. It is the first in a series of about 30 paintings that he created.

  B. It may have been painted by one of his students.

  C. it was in his possession until his death in 1675.

  D. It has the same setting as several other works of his.

  7. The word "faithful" in the passage is closest in meaning to

  A. unusual

  B. extensive

  C. exact

  D. historical

  8. Why does the author provide the information that 'When one of Vermeer's paintings was X-rayed, it did not have any preliminary sketches on the canvas beneath the paint, but rather the complete image drawn in black and white without any trial sketches"?

  A. To provide an example of a way to learn about the practices of artists who did not keep good records

  B. To emphasize Vermeer’s confidence and skill as an artist

  C. To provide evidence that Vermeer may have traced the image using a camera obscura

  D. To argue that Vermeer did his preliminary sketching on paper, rather than on canvas

  9. According to paragraph 3, Hockney believes that all of the following indicate use of a camera obscura EXCEPT

  A. very detailed, realistic work

  B. increased contrast between light and dark

  C. oversimplification of forms when the image is traced

  D. complicated foreshortening of objects

  10. The word "ntricate” in the passage is closest in meaning to

  A. surprising

  B. complex

  C. beautiful

  D. clear

  11. The word “diminish" in the passage is closest in meaning to

  A. reduce

  B. affect

  C. reflect

  D. determine

  12. According to paragraph 4，what does Hockney argue about the use of the camera obscura in producing art?

  A. Works produced using a camera obscura do not deserve as much respect as those produced without it.

  B. The camera obscura was probably used primarily as a training device, rather than used in producing finished works.

  C. Use of the camera obscura by Western art’s great masters was probably relatively rare.

  D. While the use of the camera obscura may have helped artists, they still needed to have significant artistic ability.

  13. Look at the four squares [■] that indicate where the following sentence could be added to the passage.

  All these developments helped artists to create accurate images of objects, people, and scenes.

  Where would the sentence best fit? Click on a square [■] to add the sentence to the passage

  14. Directions: An introductory sentence for a brief summary of the passage is provided below. Complete the summary by selecting the THREE answer choices that express the most important ideas in the passage. Some sentences do not belong in the summary because they express ideas that are not presented in the passage or are minor ideas in the passage. This question is worth 2 points.

  Drag your answer choices to the spaces where they belong. To remove an answer choice, click on it.

  To review the passage, click VIEW TEXT

  The camera obscura is a darkened enclosure into which light is admitted through a lens in a small hole

  A. Evidence that the use of the camera obscura has long been known is provided by its description in many sources, including works dating back to Chinese writers from the fifth century B.C.

  B. Some historians who have studied paintings by Western masters have found clues indicating that the masters may have secretly used the camera obscura in their works.

  C. It is now widely believed that the use of the camera obscura led to the development of a style of photographic realism in Western art.

  D. The camera obscura was most widely used by artists in Seventeenth-century Netherlands, a period when art and science thrived

  E. The unique features of Vermeer’s The Art of Painting make it unlikely that it was made with a camera obscura, as opposed to his other works.

  F. The artist David Hockney has speculated that artists probably combined the use of the camera A with their own original observations from life.

  3. Seagrasses

  Many areas of the shallow sea bottom are covered with a lush growth of aquatic flowering plants adapted to live submerged in seawater. These plants are collectively called seagrasses. Seagrass beds are strongly influenced by several physical factors. The most significant is water motion: currents and waves. Since seagrass systems exist in both sheltered and relatively open areas, they are subject to differing amounts of water motion. For any given seagrass system, however, the water motion is relatively constant. Seagrass meadows in relatively turbulent waters tend to form a mosaic of individual mounds, whereas meadows in relatively calm waters tend to form flat, extensive carpets. The seagrass beds, in turn, dampen wave action, particularly if the blades reach the water surface. This damping effect can be significant to the point where just one meter into a seagrass bed the wave motion can be reduced to zero. Currents are also slowed as they move into the bed.

  The slowing of wave action and currents means that seagrass beds tend to accumulate sediment. However, this is not universal and depends on the currents under which the bed exists. Seagrass beds under the influence of strong currents tend to have many of the lighter particles, including seagrass debris, moved out, whereas beds in weak current areas accumulate lighter detrital material. It is interesting that temperate seagrass beds accumulate sediments from sources outside the beds, whereas tropical seagrass beds derive most of their sediments from within.

  Since most seagrass systems are depositional environments, they eventually accumulate organic material that leads to the creation of fine-grained sediments with a much higher organic content than that of the surrounding unvegetated areas. This accumulation, in turn, reduces the water movement and the oxygen supply. The high rate of metabolism (the processing of energy for survival) of the microorganisms in the sediments causes sediments to be anaerobic (without oxygen) below the first few millimeters. According to ecologist J. W. Kenworthy, anaerobic processes of the microorganisms in the sediment are an important mechanism for regenerating and recycling nutrients and carbon, ensuring the high rates of productivity—that is, the amount of organic material produced-that are measured in those beds. In contrast to other productivity in the ocean, which is confined to various species of algae and bacteria dependent on nutrient concentrations in the water column, seagrasses are rooted plants that absorb nutrients from the sediment or substrate. They are, therefore, capable of recycling nutrients into the ecosystem that

  would otherwise be trapped in the bottom and rendered unavailable.

  Other physical factors that have an effect on seagrass beds include light, temperature, and desiccation (drying out). For example, water depth and turbidity (density of particles in the water) together or separately control the amount of light available to the plants and the depth to which the seagrasses may extend. Although marine botanist W. A. Setchell suggested early on that temperature was critical to the growth and reproduction of eelgrass, it has since been shown that this particularly widespread seagrass grows and reproduces at temperatures between 2 and 4 degrees Celsius in the Arctic and at temperatures up to 28 degrees Celsius on the northeastern coast of the United States. Still, extreme temperatures, in combination with other factors, may have dramatic detrimental effects. For example, in areas of the cold North Atlantic, ice may form in winter. Researchers Robertson and Mann note that when the ice begins to break up, the wind and tides may move the ice around, scouring the bottom and uprooting the eelgrass. In contrast, at the southern end of the eelgrass range, on the southeastern coast of the United States, temperatures over 30 degrees Celsius in summer cause excessive mortality. Seagrass beds also decline if they are subjected to too much exposure to the air. The effect of desiccation is often difficult to separate from the effect of temperature. Most seagrass beds seem tolerant of considerable changes in salinity (salt levels) and can be found in brackish (somewhat salty) waters as well as in full- strength seawater.

  1. According to paragraph 1, which of the following is true about seagrasses in calm ocean waters? A. They will not survive for very long without the nutrients brought In by fast-moving waters.

  B. They tend to form beds covering large areas along the ocean floor.

  C. They usually are arranged in separate mounds.

  D. They grow more slowly than do seagrasses in fast-moving waters.

  2. According to paragraph 1, which of the following is MOST likely to describe a bed in which seagrasses reach the surface of the water?

  A. The water is almost completely still.

  B. The bed often has major damage from strong waves or currents.

  C. The bed is generally no more than one square meter in size.

  D. Grasses form a mosaic of individual mounds.

  3. Which of the sentences below best expresses the essential information in the highlighted sentence in the passage? Incorrect choices change the meaning in important ways or leave out essential information.

  A. Light particles and debris collect in some seagrass beds, but are washed out of those affected by strong currents.

  B. Seagrass beds under the influence of strong currents tend to accumulate many of the lighter particles from other beds

  C. The strength of the currents determines how quickly accumulated seagrass debris is moved out of the beds.

  D. Seagrass debris and other light particles are often moved from areas of strong currents to areas of weak currents.

  4. The word "derive" in the passage is closest in meaning to

  A. maintain

  B. expel

  C. obtain

  D. enrich

  5. According to paragraph 3, which of the following does NOT accurately describe the sediments that collect in seagrass beds?

  A. Fine-grained

  B. Only a few millimeters deep

  C. Low in oxygen

  D. Rich in organic matter

  6. The word "confined" in the passage is closest in meaning to

  A. related

  B. limited

  C. relevant

  D. helpful

  7. According to paragraph 3，how do seagrasses affect the nutrient supply in the ecosystem?

  A. Because of their high rate of metabolism, they consume a large percentage of the available nutrients.

  B. They attract various species of algae and bacteria that produce high nutrient concentrations in the water column.

  C. They take up carbon and other nutrients trapped on the sea bottom and bring them back into use.

  D. Through anaerobic processes at their roots, they produce a very nutrient-rich sediment.

  8. It can be inferred from paragraph 4 that the reason seagrasses do not grow in very deep water is that

  A. they cannot handle intense water pressure

  B. deep water is too cold

  C. they would not get enough light

  D. deep water is too salty

  9. The word “detrimental'’ in the passage is closest in meaning to

  A. harmful

  B. significant

  C. unexpected

  D. distinct

  10. The word “detrimental'’ in the passage is closest in meaning to

  A. harmful

  B. significant

  C. unexpected

  D. distinct

  11. Paragraph 4 suggests that which of the following would be the LEAST likely to cause major damage to eelgrass and other common seagrasses?

  A. Factors related to extreme temperatures

  B. Exposure to air

  C. Major changes in salinity

  D. The movement of ice on the seafloor

  12. The phrase “tolerant of’ in the passage is closest in meaning to

  A. unused to

  B. strongly affected by

  C. protected from

  D. able to withstand

  13. Look at the four squares [■] that indicate where the following sentence could be added to the passage.

  Seagrasses grow together in dense patches, or beds, with as many as 4,000 blades per square meter.

  Where would the sentence best fit? Click on a square [■] to add the sentence to the passage

  14. Look at the four squares [■] that indicate where the following sentence could be added to the passage.

  Seagrasses grow together in dense patches, or beds, with as many as 4,000 blades per square meter.

  Where would the sentence best fit? Click on a square [■] to add the sentence to the passage

  Drag your answer choices to the spaces where they belong. To remove an answer choice, click on it.

  To review the passage, click VIEW TEXT

  Seagrasses are aquatic flowering plants that grow in either sheltered or open areas of the sea

  A. Seagrass beds are influenced by several physical factors, the most significant being the stability of the sea bottom, which must anchor them against the currents.

  B. Because they slow currents and waves, seagrass beds collect deposits of rich organic sediments, which are home to many anaerobic microorganisms.

  C. Unlike sea organisms that depend on the water column for their productivity, seagrasses ensure high rates of productivity by taking nutrients from ocean floor sediment.

  D. Sediments in seagrass beds vary by region, with temperate beds accumulating sediments from within, and tropical beds collecting sediments from without.

  E. Seagrasses under weak currents tend to have higher rates of metabolism than those under strong currents, perhaps because of differences in oxygen levels.

  F. Although seagrasses survive in temperatures ranging from 2 to 28 degrees Celsius, more extreme temperatures can damage them, as can desiccation and lack of light.

  托福TPO阅读44文本+题目+答案

  阅读一:1-5.BDACC 6-10.DAADB 11-13.CAA 14.AEF

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  阅读三:1-5.BAACB 6-10.BCCDA 11-13.CDB 14.BCF

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